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Mature Red Deer (Cervus elaphus) usually stay in single-sex groups for most of the year. During the mating ritual, called the rut, mature stags compete for the attentions of the hinds and will then try to defend hinds that they attract. Rival stags challenge opponents by belling and walking in parallel. This allows combatants to assess each other's antlers, body size and fighting prowess. If neither stag backs down, a clash of antlers can occur, and stags sometimes sustain serious injuries. Dominant stags follow groups of hinds during the rut, from August into early winter. The stags may have as many as 20 hinds to keep from other less attractive males. Only mature stags hold harems (groups of hinds) and breeding success peaks at about 8 years of age. Stags 2â€“4 years old rarely hold harems and spend most of the rut on the periphery of larger harems, as do stags over 11 years old. Young and old stags that do acquire a harem hold it later in the breeding season than those stags in their prime. Harem holding stags rarely feed and lose up to 20% of their body weight. Stags that enter the rut in poor condition are less likely to make it through to the peak conception period. Male European Red Deer have a distinctive "roar" during the rut, which is an adaptation to forested environments, as opposed to male Wapiti (or American Elk) which "bugle" during the rut in adaptation to open environments. The male deer roars to keep his harem of females together. The females are initially attracted to those males that both roar most often and have the loudest roar call. Males also use the roar call when competing with other males for females during the rut, and along with other forms of posturing and antler fights, is a method used by the males to establish dominance. Roaring is most common during the early dawn and late evening, which is also when the crepuscular deer are most active in general.
Breeding, gestation and lifespan
Red Deer mating patterns usually involve a dozen or more mating attempts before the first successful one. There may be several more matings before the stag will seek out another mate in his harem. Females in their second autumn can produce one and very rarely two offspring per year. The gestation period is 240 and 262 days and the offspring weigh about 15 kg (33 lb). After two weeks, calves are able to join the herd and are fully weaned after two months. Female offspring outnumber male offspring more than two to one and all Red Deer calves are born spotted, as is common with many deer species, and lose their spots by the end of summer. However, as in many species of Old World Deer, some adults do retain a few spots on the backs of their summer coats. The offspring will remain with their mothers for almost one full year, leaving around the time that the next season offspring are produced. The gestation period is the same for all subspecies. Red Deer live up to over 20 years in captivity and in the wild they average 10 to 13 years,, though some subspecies with less predation pressure average 15 years.
The Red Deer is one of the largest deer species. It is a ruminant, eating its food in two stages and having an even number of toes on each hoof, like camels, goats and cattle. European Red Deer have a relatively long tail compared to their Asian and North American relatives. There are subtle differences in appearance between the various subspecies of Red Deer primarily in size and antlers, with the smallest being the Corsican Red Deer found on the islands of Corsica and Sardinia and the largest being the Caspian red deer (or maral) of Asia Minor and the Caucasus Region to the west of the Caspian Sea. The deer of Central and Western Europe vary greatly in size with some of the largest deer found in the Carpathian Mountains in Central Europe. West European Red Deer historically, grew to large size given ample food supply (including peoples' crops), and descendants of introduced populations living in New Zealand and Argentina have grown quite large in size and antlers. Large Red Deer stags, like the Caspian Red Deer or those of the Carpathian Mountains may rival the Wapiti in size. Female Red Deer are much smaller than their male counterparts. Generally, the male (stag or hart) Red Deer is typically 175 to 230 cm (69 to 91 in) long and weighs 160 to 240 kg (350 to 530 lb); the female is 160 to 210 cm (63 to 83 in) long and weighs 120 to 170 kg (260 to 370 lb). The tail adds another 12 to 19 cm (4.7 to 7.5 in) and shoulder height is about 105 to 120 cm (41 to 47 in). Size varies in different subspecies with the largest, the huge but small-antlered deer of the Carpathian Mountains (C. e. elaphus), weighing up to 500 kg (1,100 lb). At the other end of the scale, the Corsican Red Deer (C. e. corsicanus) weighs about 80 to 100 kg (180 to 220 lb), although Red Deer in poor habitats can weigh as little as 53 to 112 kg (120 to 250 lb). European Red Deer tend to be reddish-brown in their summer coats. The males of many subspecies also grow a short neck mane ("mane" of hair around their necks) during the autumn. The male deer of the British Isles and Norway tend to have the thickest and most noticeable neck manes. Male Caspian Red Deer (Cervus elaphus maral) and Spanish Red Deer (Cervus elaphus hispanicus) do not carry neck manes. Male deer of all subspecies, however, tend to have stronger and thicker neck muscles than female deer, which may give them an appearance of having neck manes. Red Deer hinds (females) do not have neck manes. The European Red Deer is adapted to a woodland environment. Only the stags have antlers which start growing in the spring and are shed each year, usually at the end of winter. Antlers are made of bone which can grow at a rate of 2.5 cm (1.0 in) a day. A soft covering known as velvet helps to protect newly forming antlers in the spring. European red deer antlers are distinctive in being rather straight and rugose, with the fourth and fifth tines forming a "crown" or "cup" in larger males. Any tines in excess of the fourth and fifth tine will grow radially from the "cup". "Cups" are generally absent in the antlers of smaller red deer such as Corsican Red Deer. West European Red Deer antlers feature bez (second) tines that are either absent or smaller than the brow tine. However, bez tines occur frequently in Norwegian Red Deer. Antlers of Caspian Red Deer carry large bez (second) tines and form less-developed "cups" than West European red deer. A stag can (exceptionally) have smooth antlers, and is then known as a switch. Similarly, a stag that doesn't grow antlers is a hummel. The antlers are testosterone-driven and as the stag's testosterone levels drop in the autumn, the velvet is shed and the antlers stop growing.. Red Deer produce no testosterone in their bodies while they are growing antler.[clarification needed] With the approach of autumn, the antler begin to calcify and the stags testosterone production builds for the approaching rut (mating season). During the autumn, all Red Deer subspecies grow a thicker coat of hair which helps to insulate them during the winter. Autumn is also when some of the stags grow their neck manes. It is in the autumn/winter coat that most subspecies are most distinct. The Caspian Red Deer's winter coat is greyer and has a larger and more distinguished light rump-patch (like Elk and some Central Asian Red Deer) compared to the West European Red Deer which has more of a greyish-brown coat with a darker yellowish rump patch in the winter. By the time summer begins, the heavy winter coat has been shed; the animals are known to rub against trees and other objects to help remove hair from their bodies. Red Deer have different colouration based on the seasons and types of habitats, with grey or lighter colouration prevalent in the winter and a more reddish and darker coat in the summer. Most European Red Deer wear a reddish-brown summer coat, and some individuals may have a few spots on the backs of their summer coats.
Distribution and habitat
Cervus genus ancestors of Red Deer first appear in fossil records 12 million years ago during the Pliocene in Eurasia.  An extinct species, known as the Irish Elk (Megaloceros) was not related to the red deer but to the fallow deer, was the largest member of the deer family known from the fossil record. The European Red Deer is one of the largest game animals found in Southwestern Asia (Asia Minor and Caucasus Regions), North Africa and Europe. The Red Deer is the largest non-domesticated mammal still existing in some European countries such as the United Kingdom and Ireland. The Barbary stag (which resembles the West European Red Deer) is the only member of the deer family that is represented in Africa, with population centred in the northwestern region of the continent in the Atlas Mountains. As of the mid 1990s, Morocco, Tunisia and Algeria were the only African countries known to have Red Deer. In the UK there are indigenous populations in Scotland, the Lake District and the South West of England (principally on Exmoor). Not all of these are of entirely pure bloodlines as some of these populations have been supplemented with deliberate releases of deer from parks like Warnham or Woburn Abbey in an attempt to increase antler sizes and body weights, and particularly in Scotland there has been extensive hybridisation with the closely related Sika Deer. There are several other populations that have originated either with carted deer kept for stag hunts being left out at the end of the hunt,[clarification needed] escapes from deer farms or deliberate releases. Carted deer were kept by stag hunts with no wild red deer in the locality and were normally recaptured after the hunt and used again. Sometimes they escaped and formed the basis of a new population, the red deer based on Thetford Forest in Norfolk came from this source. There are now substantial red deer herds that originated from escapes or deliberate releases in the New Forest, the Peak District, Suffolk, Brecon Beacons and West Yorkshire as well as many other smaller populations scattered throughout England, and they are all generally increasing in numbers and range. In New Zealand, and to a much lesser degree in Australia, there are only a couple of small herds left of the original deer brought over for attempted breeding. Most Red deer in Australia are on hunting preserves. In New Zealand introduced Red Deer have adapted well and are widely hunted on both islands, these herds originated from deer bought from Scotland (Invermark) or one of the deer parks in England, principally Warnham, Woburn Abbey or Windsor Great Park.[clarification needed] New Zealand red deer produce very large antlers and are regarded as amongst the best in the world by hunters. Along with the other introduced deer species they are however officially regarded as a noxious pest and have at times been heavily culled using professional hunters working with helicopters, or even poisoned. Red Deer populations in Africa and southern Europe are generally declining. In Argentina, where the Red Deer has had a potential adverse impact on native animal species, the International Union for Conservation of Nature and Natural Resources has labelled the animal as one of the world's 100 worst invaders.
Red Deer in Europe generally spend their winters in lower altitudes and more wooded terrain. During the summer, they migrate to higher elevations where food supplies are greater for the calving season.
Protection from predators
Male Red Deer retain their antlers for more than half the year and are less gregarious and less likely to group with other males when they have antlers. The antlers provide self-defence as does a strong front-leg kicking action which is performed by both sexes when attacked. Once the antlers are shed, stags tend to form bachelor groups which allow them to cooperatively work together. Herds tend to have one or more members watching for potential danger while the remaining members eat and rest. After the rut, females form large herds of up to 50 individuals. The newborn calves are kept close to the hinds by a series of vocalizations between the two, and larger nurseries have an ongoing and constant chatter during the daytime hours. When approached by predators, the largest and most robust females may make a stand, using their front legs to kick at their attackers. Guttural grunts and posturing is used with all but the most determined of predators with great effectiveness. Aside from humans and domestic dogs, the Wolf is probably the most dangerous predator that most European Red Deer encounter. Occasionally, the Brown bear will predate on European Red Deer as well. Eurasian Lynx and wild boars sometimes prey on the calves. The leopard in Asia Minor (now extinct) probably preyed on East European Red Deer. Both Barbary Lion and Barbary Leopard probably once preyed on Atlas stags in the Atlas Mountains, although Barbary Lion is now extinct in the wild, and Barbary Leopard either very rare or extinct.
Red Deer in folklore
Red Deer are widely depicted in cave art and are found throughout European caves, with some of the artwork dating from as early as 40,000 years ago, during the Upper Paleolithic. Siberian cave art from the Neolithic of 7,000 years ago has abundant depictions of Red Deer, including what can be described as spiritual artwork, indicating the importance of this mammal to the peoples of that region (Note: these animals were most likely Wapiti (Cervus canadensis) in Siberia, not Red Deer). Red deer are also often depicted on Pictish stones (c.550-850 AD), from the early medieval period in Scotland, usually as prey animals for human or animal predators.
Red Deer products
Red Deer are held in captivity for a variety of reasons. The meat of the deer, called venison, is not generally harvested for human consumption on a large scale, though speciality restaurants seasonally offer venison which is widely considered to be both flavourful and nutritious. Venison is higher in protein and lower in fat than either beef or chicken. In some countries in central Asia, elk is still hunted as a primary source of meat. The red deer can produce 10 to 15 kg (22 to 33 lb) of antler velvet annually. On ranches in New Zealand, China, Siberia, and elsewhere this velvet is collected and sold to markets in East Asia, where it is used for holistic medicines, with South Korea being the primary consumer. In Russia, a medication produced from antler velvet is sold under the brand name Pantokrin (Russian: ÐŸÐ°Ð½Ñ‚Ð¾ÐºÑ€Ð¸ÌÐ½; Latin: Pantocrinum). The antlers themselves are also believed by East Asians to have medicinal purposes and are often ground up and used in small quantities. Historically, related deer species such as Central Asian Red Deer, Wapiti, Thorold's Deer, and Sika Deer have been been reared on deer farms in Central and Eastern Asia by Han Chinese, Turkic peoples, Tungusic peoples, Mongolians, and Koreans. In modern times, Western countries such as New Zealand and United States have taken to farming European Red Deer for similar purposes. Deer antlers are also highly sought after worldwide for decorative purposes and have been used for artwork, furniture and other novelty items.
Additionally there are some central asiatic subspecies (Tarim group, including Bactrian deer and Yarkand deer), which are geographically isolated from Wapiti and western Red Deer by the Takla Makan and the Pamir Mountains. They appear to represent a primordial subgroup, genetically more related to the Red Deer than to the Wapiti. It remains unclear which clade the Kashmir stag belongs in, though it, in terms of zoogeography, is most likely to belong in the central Asian group. The International Union for Conservation of Nature and Natural Resources originally listed nine subspecies of Red Deer (Cervus elaphus): three as endangered, one as vulnerable, one as near threatened, and four without enough data to give a category ("Data Deficient"). The species as a whole, however, is listed as least concern.. However, this was based on the traditional classification of Red Deer as one species (Cervus elaphus), including the Elk. Listed below are the subspecies of Red Deer (Cervus elaphus), including the primordial subgroup from central Asia.
Until recently biologists considered that Red Deer and Wapiti (or Elk) are the same species forming, a continuous distribution throughout temperate Eurasia and North America. This belief was based largely on the fully fertile hybrids that can be produced under captive conditions. However, recent DNA studies conducted on hundreds of samples from Red Deer and Elk subspecies concluded that there are no more than 9 distinct subspecies of Red Deer and Wapiti and that they fall into two separate species: the Red Deer from Europe, western Asia and North Africa, and the Wapiti or Elk from Northern and Eastern Asia and North America. Surprisingly, from DNA evidence the Elk appear more closely related to Sika Deer and to Thorold's deer than to Red Deer.
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